Tag Archives: Red-Tailed Hawk

A Somewhat Disappointing Excursion, 05-23-19

Feeling tired and stressed, I decided to try making the drive to the Sacramento National Wildlife Refuge. Rather than being a therapeutic and relaxing time, however, I was faced with irritation after irritation, so the drive actually left me feeling more stressed and exhausted than I was before I started. *Heavy sigh*

CLICK HERE for the album of photos.

Traffic wasn’t too terribly terrible, but it seemed like I hit every freaking red light there was, and the monster semi’s on the highway were “always” in front of me like giant snails. When I got to the refuge, I could see Bank Swallows collecting mud in the slough near the front gate, so I stopped to take some photos – and, of course, the birds wouldn’t cooperate, and two employees pulled up in their cars behind me and honked at me. Grrrrr. The rest of the day kind of went like that: I saw Gallinules, but they ducked into the tules before I could get a photo; I could hear Bitterns giving their “pumperlunk” calls close by, but couldn’t see them; it was windy, so most of the birds were hunkered down near the ground or deep in the trees; where there would normally be dozens of Marsh Wrens around, I saw only two deep in the tules; the dragonflies hadn’t come up from the water yet; what damselflies there were around were the tiniest ones that are like trying to photograph a strand of hair; there weren’t any of the Clark’s or Western Grebes that I was expecting to be out there by now; I couldn’t even get shots of ground squirrels… It was just one frustration after another. By the time I left, I had a splitting headache and just wanted to ram someone with my car. ((I didn’t though.)) *Heavy sigh-2*

Species List:

  1. American Avocet, Recurvirostra americana,
  2. American Bittern, Botaurus lentiginosus,
  3. American Bullfrog, Lithobates catesbeianus,
  4. American White Pelican, Pelecanus erythrorhynchos,
  5. Bank Swallow, Riparia riparia,
  6. Black Phoebe, Sayornis nigricans,
  7. Black-Tailed Jackrabbit, Lepus californicus,
  8. Blessed Milk Thistle, Silybum marianum,
  9. Cabbage White Butterfly, Pieris rapae,
  10. California Dock, Rumex californicus,
  11. California Milkweed, Asclepias californica,
  12. Canada Goose, Branta canadensis,
  13. Cinnamon Teal, Anas cyanoptera,
  14. Columbian Black-Tailed Deer, Odocoileus hemionus columbianus,
  15. Common Gallinule, Gallinula galeata,
  16. Convergent Lady Beetle, Hippodamia convergens,
  17. Desert Cottontail, Sylvilagus audubonii,
  18. Double-Crested Cormorant, Phalacrocorax auritus,
  19. Downingia, Downingia sp.,
  20. English Lawn Daisy, Bellis perennis,
  21. Eurasian Collared Dove, Streptopelia decaocto,
  22. Familiar Bluet Damselfly, Enallagma civile,
  23. Goodding’s Willow, Salix gooddingii,
  24. Great Blue Heron, Ardea herodias,
  25. Great Egret, Ardea alba,
  26. Greater White-Fronted Goose, Anser albifrons,
  27. Italian Thistle, Carduus pycnocephalus,
  28. Mallard, Anas platyrhynchos,
  29. Marsh Wren, Cistothorus palustris,
  30. Mourning Dove, Zenaida macroura,
  31. Mute Swan, Cygnus olor,
  32. Northern Pintail, Anas acuta,
  33. Northern Shoveler, Anas clypeata,
  34. Painted Lady Butterfly, Vanessa cardui,
  35. Poison Hemlock, Conium maculatum,
  36. Red-Tailed Hawk, Buteo jamaicensis,
  37. Red-Winged Blackbird, Agelaius phoeniceus,
  38. Ring-Necked Pheasant, Phasianus colchicus,
  39. Short-tailed ichneumon wasps, Ophion sp.,
  40. Snowy Egret, Egretta thula,
  41. Song Sparrow, Melospiza melodia,
  42. Tule, Schoenoplectus acutus,
  43. Variegated Meadowhawk Dragonfly, Sympetrum corruptum,
  44. Western Fence Lizard, Sceloporus occidentalis,
  45. Western Kingbird, Tyrannus verticalis,
  46. Wild Teasel, Dipsacus fullonum,

A Lot of Little Moments, 05-14-19

I got up at 5:00 this morning and took my time getting ready to head out to the Effie Yeaw Nature Preserve for my trail-walking thing there. It was cool for most of the day – a rainstorm is supposed to move in tomorrow – so it was perfect walking weather. I was joined by fellow volunteer Mary Messenger – the Other Mary.

We saw the usual suspects like deer, House Wrens, and suchlike. One humorous encounter was with a European Staring who had its adult feathers in but was acting like a brat. It was sitting in a tree just opposite where I had seen the fledglings poking their heads out of the nesting cavity last week. It was making a lot of very loud squawks and peeps, and flapping its wings trying to get attention. Might have also been a female looking for a mate to come feed her. Whichever.  She was so loud and so animated; you couldn’t miss her.

Another funny moment was walking in on a pair of Fox Squirrels who I think were making out.  Hah! Get a room, you guys!

CLICK HERE for the album of photos.

I saw another Starling in another part of the preserve that was taking twigs OUT of her nesting cavity, which I thought was weird.  And we saw a male Mourning Dove picking up bits of grass and carrying them to his mate in a tree off the trail.  A House Wren was carrying food to his babies… Everyone was moving stuff around.

I also saw a pair of Wood Ducks. They flew into a tree overhead, and then the female flew to an adjacent tree and “disappeared”. She flew out to the first tree next to the male, then flew back to the other tree and disappeared again. I tried to see where she was hiding out and assumed she might have had a nest in a cavity in the tree, but I just couldn’t see her. Then she flew out one more time, and this time she had something orange and fuzzy in her bill. She flew off with it, and the male followed her. I couldn’t tell what it was, really, and wasn’t able to get any photos of it, but I think she was retrieving a duckling that was refusing to come down from the nest! I’d never seen or heard of anything like that; it was kind of amazing.

And we caught a glimpse of a young coyote. He came out onto the trail in front of us with a short growl and then ran off into the high grass where we lost sight of him. He was pretty small, probably a teenager. After we saw him, we kept an eye out for mom and dad; they usually travel in a pack when the pups are young.

We walked for about 4 hours, which is pretty much the limit for both of us, and I headed back home.

Species List:

  1. Acorn Woodpecker, Melanerpes formicivorus,
  2. American Bullfrog, Lithobates catesbeianus,
  3. American Robin, Turdus migratorius,
  4. Ash-Throated Flycatcher, Myiarchus cinerascens,
  5. Asian Lady Beetle, Harmonia axyridis,
  6. Black Grosbeak, Pheucticus melanocephalus,
  7. Black Phoebe, Sayornis nigricans,
  8. Blessed Milk Thistle, Silybum marianum,
  9. Blue Elderberry, Sambucus cerulea,
  10. California Buckeye Chestnut Tree, Aesculus californica,
  11. California Pipevine Swallowtail Butterfly, Battus philenor hirsuta,
  12. California Pipevine, Dutchman’s Pipe, Aristolochia californica,
  13. Coffeeberry, Frangula californica,
  14. Columbian Black-Tailed Deer, Odocoileus hemionus columbianus,
  15. Coyote Brush Bud Gall Midge, Rhopalomyia californica,
  16. Coyote, Canis latrans,
  17. Darkling Beetle, Eleodes dentipes,
  18. Desert Cottontail, Sylvilagus audubonii,
  19. Eastern Fox Squirrel, Sciurus niger,
  20. Elegant Clarkia, Clarkia unguiculata,
  21. European Starling, Sturnus vulgaris,
  22. Goldwire, Hypericum concinnum,
  23. Great Egret, Ardea alba,
  24. Greater Periwinkle, Vinca major,
  25. House Wren, Troglodytes aedon,
  26. Italian Thistle, Carduus pycnocephalus,
  27. Katydid nymph, Microcentrum rhombifolium,
  28. Miniature Lupine, Lupinus bicolor,
  29. Mourning Dove, Zenaida macroura,
  30. Oak Apple Wasp Galls, Andricus quercuscalifornicus,
  31. Oak Titmouse, Baeolophus inornatus,
  32. Olive Tree, Olea europaea,
  33. Pacific Pond Turtle, Western Pond Turtle, Actinemys marmorata,
  34. Painted Lady Butterfly, Vanessa cardui,
  35. Pinacate Beetle, Eleodes pimelioides,
  36. Red Harvester Ant, Pogonomyrmex barbatus,
  37. Red-Shouldered Hawk, Buteo lineatus,
  38. Red-Tailed Hawk, Buteo jamaicensis,
  39. Rusty Tussock Moth caterpillar, Orgyia antiqua,
  40. Spittle Bug, Meadow Spittlebug, Philaenus spumarius,
  41. Spotted Towhee, Pipilo maculatus,
  42. Swainson’s Hawk, Orion, Buteo swainsoni,
  43. Tree Swallow, Tachycineta bicolor,
  44. Wavy-Leafed Soap Plant, Soaproot, Chlorogalum pomeridianum,
  45. Western Gray Squirrel, Sciurus griseus,
  46. Winter Vetch, Vicia villosa,
  47. Wood Duck, Aix sponsa,

At the Sacramento and Colusa Wildlife Refuges, 05-06-19

Certified California Naturalist, Roxanne Moger and I went out to the Sacramento National Wildlife Refuge around 6:00 am this morning. The weather was beautiful today, but there wasn’t a lot to see at the preserve. We’re kind of in between seasons, so the large flocks of birds have all left, but the insects (like dragonflies, damselflies, and orb-weaver spiders) haven’t arrived yet. We did see swarms of Painted Lady butterflies and some Cabbage White, but none of the other species normally seen there in the late spring/early summer.

Roxanne did all the driving, but we stopped a couple of time to get out of the car and walk parts of the trail or explore the boundaries of the park-and-stretch areas. Some of the vernal pools on the site were in bloom: all golden yellow and purple with Goldfields and Downingia. Just beautiful.

Although the species list at the end of the day wasn’t as long or as varied as I’d like, I did like the fact that I saw a few things I’d never seen before and learned more about some species than I’d known before. That’s what really makes these outings fun.

A large fly landed on the passenger side mirror of the car partway through the auto-tour and at first I thought it was a Robber Fly (those guys are pretty big). We were both intrigued by the fly’s huge goggle-like eyes and his tenaciousness.  He held onto that mirror for quite a long time.

When I got home, I looked up the fly to see if I could find its scientific name and found that it was actually a male Striped (or Lined) Horsefly, Tabanus lineola. I’d never seen one before. And, of course, once I find something new to me, I have to research it more.  Never having encountered a Horsefly before, I was surprised to learn that although the males drink nectar, the females drink blood (usually from large mammals like livestock). She had scissor-like mouth parts that slice into the skin so she can get to the blood. This species is usually found along the east coast and Gulf of Mexico, so it was something of a surprise to find it here… assuming I got the ID correct.

We also got to see a male Red-Winged Blackbird flaring his epaulets at a female, and a pair of Brown-Headed Cowbirds performing what we thought was courtship behavior.  The two birds sat across from one another, then one ruffled its feathers, opened its wings a bit and bowed down at the other, and the other responded in kind.

Roxanne and I inferred the behavior was “courtship” but, from what I read, after I got home, what we were seeing was actually two males trying to outdo one another in a machismo contest.  Apparently, the females don’t respond well to the males’ bows, which they see as aggressive, so the males only bow to one another. (You can read more here.)  The males open their wings to one another, and to females, to show how mature they are. Juveniles have pale markings on the inside of the wings.

You can see the video snippet I took of the birds HERE.

CLICK HERE for the full album of photos.

And we found a couple of Black Phoebe nests, one stuffed full of babies who were almost fully fledged. They were stacked up, one on top of the other, with their tails sticking out over the edge of the nest. And toward the end of the auto tour route we came across a dead tree where there was a Western Meadowlark, a Red-Winged Blackbird and a tiny Song Sparrow all singing their respective songs.

We caught glimpses of American Goldfinches and Bullock’s Orioles, and hear Bitterns, but didn’t see any. One oddity was sighting a Mute Swan in the permanent wetlands area. That was odd because swans hardly ever go into the refuge, and Mute Swans are actually an invasive species. Unlike the Tundra Swans, the Mute Swans are super-aggressive and destroy the habitat they live in by ripping up water-plants from the roots.

When we were done at the Sacramento refuge, we decided to go over to the Colusa National Wildlife Refuge to show Roxanne the day-roost of the Black-Crowned Night Herons at the end of the auto-tour there. We were astonished to discover that the auto-tour there was roped off so no one could get to it… but the rope and signage was only visible AFTER you entered the refuge and started driving down the route. Stupid. They should have put the signs on the front gate or in the parking area.  A “manager” who showed up a little while after did, said that the auto-route was shut down because they were short handed and couldn’t patrol it well enough. Sad.

So, we didn’t do that tour and instead walked around a little bit in the native flower garden they have near the restroom facility.  Along part of a path near the garden, Roxanne found some galls we’d never seen before: the gall of the Elm Balloon-Gall Aphid, Eriosoma lanuginosum.

According to what I’ve read, the galls are initiated by a “fundatrix”, a parthenogenetic female aphid whose presence causes an extreme enlargement of the soft cell tissue on one side of the leaf. The galls are hollow and feel rubbery. They start out green and are covered with fine white hairs (which we saw) and turn brown as they age.

Inside the gall the fundatrix has her babies which are wingless and yellow until they mature. There can also be second generation aphids, called “alates” (usually winged individuals) which are dark green to black and wax powdered. Not all of the adults grow wings, however, and wing-growth seemed to be associated to crowding inside the galls, a short supply of food, and/or changes in the environment. Nature is so cool!

After walking around a bit, we headed back home to Sacramento and got there around 3:00 pm.

Species List:

1. American Avocet, Recurvirostra americana,
2. American Bittern, Botaurus lentiginosus,
3. American Coot, Fulica americana.
4. American Crow, Corvus brachyrhynchos,
5. American Goldfinch, Spinus tristis,
6. American Robin, Turdus migratorius,
7. Anna’s Hummingbird, Calypte anna,
8. Annual Yellow Sweet Clover, Melilotus indicus,
9. Arches Moth, Habrosyne sp.
10. Bird’s Foot Trefoil, Lotus corniculatus,
11. Black Phoebe, Sayornis nigricans,
12. Black-Tailed Jackrabbit, Lepus californicus,
13. Blessed Milk Thistle, Silybum marianum,
14. Blue Elderberry, Sambucus cerulea,
15. Bristly Oxtongue, Helminthotheca echioides,
16. Brown-Headed Cowbird, Molothrus ater,
17. Bullock’s Oriole, Icterus bullockii,
18. Bur Clover, Burr Medic, Medicago polymorpha,
19. California Flannelbush, Fremontodendron californicum,
20. California Ground Squirrel, Otospermophilus beecheyi,
21. California Poppy, Eschscholzia californica,
22. California Wild Rose, Rosa californica,
23. Canada Goose, Branta canadensis,
24. Cinnamon Teal, Anas cyanoptera,
25. Clark’s Grebe, Aechmophorus clarkia,
26. Cleveland Sage, Salvia clevelandii,
27. Common Mustard, Brassica rapa,
28. Convergent Lady Beetle, Hippodamia convergens,
29. Curly Dock, Rumex crispus,
30. Damselfly, Pacific Forktail, Ischnura cervula, (dots on thorax)
31. Damselfly, Sooty Dancer, Argia lugens, (no blue tip; rings around segments)
32. Desert Cottontail, Sylvilagus audubonii,
33. Dogtail Grass, Cynosurus echinatus,
34. Elm Balloon-Gall Aphid, Eriosoma lanuginosum,
35. Eurasian Collared Dove, Streptopelia decaocto,
36. Field Elm Tree, Smooth-Leaf Elm, Ulmus Minor,
37. Foothills Penstamon, Penstemon heterophyllus,
38. Foxtail Barley, Hordeum murinum ssp. glaucum,
39. Fremont Cottonwood Tree, Populus fremontii,
40. Fuller’s Teasel, Wild Teasel, Dipsacus fullonum,
41. Goldfields, Contra Costa Goldfields, Lasthenia conjugens,
42. Great Egret, Ardea alba,
43. Greater White-Fronted Goose, Anser albifrons,
44. Greenbottle Fly, Lucilia sericata,
45. Himalayan Blackberry, Rubus armeniacus,
46. Hood Canarygrass, Phalaris paradoxa,
47. Hoover’s Downingia, Hoover’s Calicoflower, Downingia bella,
48. House Finch, Haemorhous mexicanus,
49. House Sparrow, Passer domesticus,
50. Jimson Weed, Datura stramonium,
51. Killdeer, Charadrius vociferous,
52. Large Oxtongue Aphid, Uroleucon picridis,
53. Marsh Wren, Cistothorus palustris,
54. Mourning Dove, Zenaida macroura,
55. Mute Swan, Cygnus olor,
56. Narrowleaf Milkweed, Asclepias fascicularis,
57. Northern Paper Wasp, Polistes fuscatus,
58. Northern Pintail, Anas acuta,
59. Northern Shoveler, Anas clypeata,
60. Oleander Aphid, Aphis nerii,
61. Pacific Pond Turtle, Western Pond Turtle, Actinemys marmorata,
62. Pacific Tree Frog, Chorus Frog, Pseudacris regilla,
63. Painted Lady Butterfly, Vanessa cardui,
64. Pied-Billed Grebe, Podilymbus podiceps,
65. Plantain, Ribwort, Plantago lanceolata,
66. Poison Hemlock, Conium maculatum,
67. Purple Finch, Haemorhous purpureus,
68. Purple Needle Grass, Stipa pulchra,
69. Rabbit Tail Grass,Hare’s Tail Grass, Lagurus ovatus,
70. Red-Tailed Hawk, Buteo jamaicensis,
71. Red-Winged Blackbird, Agelaius phoeniceus,
72. Rush, California Bulrush, Schoenoplectus californicus,
73. Sedge, Pennsylvania Sedge, Carex pensylvanica
74. Showy Milkweed, Asclepias speciose,
75. Silverpuff, Microseris acuminata,
76. Snow Goose, Chen caerulescens,
77. Song Sparrow, Melospiza melodia,
78. Striped Horsefly, Tabanus lineola,
79. Tule, Schoenoplectus acutus,
80. Turkey Vulture, Cathartes aura,
81. Valley Popcorn Flower, Plagiobothrys canescens,
82. Varied Carpet Beetle, Anthrenus verbasci,
83. Variegated Meadowhawk Dragonfly, Sympetrum corruptum,
84. Western Fence Lizard, Sceloporus occidentalis,
85. Western Kingbird, Tyrannus verticalis,
86. Western Meadowlark, Sturnella neglecta,
87. Willow, Gooding’s Willow, Salix gooddingii

Interesting Turkeys on the Naturalist Walk, 04-09-19

Up at 6:00 this morning, and then headed out to the Effie Yeaw Nature Preserve to do my trail walking thing – along with three of my naturalist students.  The weather was lovely, so the critters were out and moving around, and the wildflowers were really starting to open up.

We got to see a tiny female American Kestrel chase off a large Cooper’s Hawk, saw the young buck with a broken nose browsing with some of the does, and saw a yard-long gopher snake leave the side of the trail and rush through the grass like water.

We also came across a female Wild Turkey sitting on the ground next to a brush pile occupied by some California Ground Squirrels. Everything I’d read about the turkeys indicated that they nested in low branches of trees or ON brush piles, so even though she looked pretty settled on the ground I assumed she was just taking a dirt bath (which the turkeys often do to control mites).  I walked up to her, slowly, and she eventually stood up and walked away from where she’d been sitting: a bare, shallow patch of dirt.

When I got home, I did some more research on the turkey and found this: “…Wild Turkeys nest on the ground in dead leaves at the bases of trees, under brush piles or thick shrubbery, or occasionally in open hayfields. The female scratches a shallow depression in the soil, about 1 inch deep, 8–11 inches wide, and 9–13 inches long…” So, this gal may have been prepping a nest site, not dirt bathing.  I’ll have to check on the spot again the next time I’m out there.

Seeing the turkeys -– including one of the leucistic females who came out of the forest like a ghost – brought on a flurry of “snood” jokes. Four women on the trail talking about the male turkeys’ accouterments. We couldn’t help ourselves. Hahahahaha!

Oh, and I also learned that although the male hierarchy changes a lot as the males challenge, defeat, and retreat from one another, the female turkeys’ hierarchy remains constant from season to season, with a dominant female overseeing all of the ladies.  How cool is that?

We didn’t see the Mourning Doves on or near their nest, and I’m afraid they may have abandoned it.  When we got to the Red-Shouldered Hawk nest on the Pond Trail, however, mama hawk was in the nest and calling out to hubby. We could hear her, but we couldn’t see her. The nest is pretty deep and it’s right over the trail so it’s hard to see into it. Suddenly, mama hawk burst out away from the nest and flew right at and over one of the students! She was able to catch a photo of the hawk as she flew over her head! Awesome!

Among the flowers we saw were Blue Dicks, Bush Lupine, California Poppies, Fringe Pod, Periwinkle, Miniature Lupine, different kinds of plantains, and a variety of tiny yellow flowers that defied identification. On one of the plants was an example of “fasciation”, wherein the flowering heads weld together.  “…Scientists aren’t sure what causes the deformity, but they believe it is probably caused by a hormonal imbalance. This imbalance may be the result of a random mutation, or it can be caused by insects, diseases or physical injury to the plant. Think of it as a random occurrence. It doesn’t spread to other plants or other parts of the same plant…”

CLICK HERE to see the album of photos.

The ladies were wonderful trail walking companions: they were excited about everything, had great questions, and lots of wonderful feedback about the naturalist course. We walked for almost 5 hours (which was way too long for me) before we all headed back home again. Deborah had come all the way from Napa to walk with me, so she had the longest drive back home (over 2 hours).

I totally overdid it and was exhausted and in pain when I got home, but it was worth it to have spent those hours with the ladies’ positive energy.  Even though I practically went back to bed when I got home, I was happy. This is just how I want to spend what time I have left.

It was a good day.

Species List:

1. Acorn Woodpecker, Melanerpes formicivorus,
2. Asian Ladybeetle, Multicolored Asian Lady Beetle, Harmonia axyridis,
3. Bay Laurel, Laurus nobilis,
4. Black Walnut, Juglans nigra,
5. Blue Dicks, Dichelostemma capitatum,
6. Blue Oak, Quercus douglasii,
7. Buck Brush, Ceanothus cuneatus,
8. Bush Lupine, Lupinus albifrons,
9. California Ground Squirrel, Otospermophilus beecheyi,
10. California Manroot, Marah fabaceus,
11. American Kestrel, Falco sparverius,
12. Black Bean Aphid, Aphis fabae.
13. Boxedler Tree, Boxelder Maple, Acer negundo.
14. Bush Lupine, Lupinus excubitus.
15. California Plantain, Plantago erecta.
16. California Pipevine Swallowtail Butterfly, Battus philenor hirsuta,
17. California Pipevine, Aristolochia californica,
18. California Poppy, Eschscholzia californica,
19. California Scrub Jay, Aphelocoma californica,
20. Columbian Black-Tailed Deer, Odocoileus hemionus columbianus,
21. Cooper’s Hawk, Accipiter cooperii,
22. Coyote Brush Midge (gall), Rhopalomyia californica,
23. Coyote Brush, Baccharis pilularis,
24. Douglas Iris, Iris douglasiana,
25. Eastern Fox Squirrel, Sciurus niger,
26. European Starling, Sturnus vulgaris,
27. Fringe Pod, Thysanocarpus curvipes ssp. elegans,
28. Green Shield Lichen,Flavoparmelia caperata,
29. House Wren, Troglodytes aedon,
30. Interior Live Oak, Quercus wislizeni,
31. Lace Lichen, Ramalina menziesii,
32. Miniature Lupine, Lupinus bicolor,
33. Mourning Dove, Zenaida macroura,
34. Nutthall’s Woodpecker, Picoides nuttallii,
35. Pacific Gopher Snake, Pituophis catenifer,
36. Painted Lady butterfly, Vanessa cardui,
37. Periwinkle, Vinca major,
38. Red-Shouldered Hawk, Buteo lineatus,
39. Red-Tailed Hawk, Buteo jamaicensis,
40. Rio Grande Wild Turkey, Meleagris gallopavo intermedia,
41. Saw-Whet Owl, Aegolius acadicus,
42. Snowy Egret, Egretta thula,
43. Spotted Towhee, Pipilo maculatus,
44. Tree Swallow, Tachycineta bicolor,
45. Turkey Vulture, Cathartes aura,
46. Valley Oak, Quercus lobata,
47. Wavy-Leaf Soap Root, Soap Plant, Chlorogalum pomeridianum,
48. Western Redbud, Cercis occidentalis,
49. Western Fence Lizard, Sceloporus occidentalis,
50. Yarrow, Common Yarrow, Achillea millefolium,